As the British Property Federation says...

… on page 16* of this useful report:

The commercial property industry is taxed directly and indirectly in a variety of ways. The direct contributions from some of these taxes – including Stamp Duty Land Tax, VAT, PAYE and National Insurance contributions – can be calculated with reasonable accuracy and are illustrated below…

Other taxes directly paid by the industry, ranging from Corporation Tax to rates on empty property and the Community Infrastructure Levy, are much harder to assess. In addition, occupiers of commercial property paid £20 billion in Business Rates, some of which, effectively, will be borne by property owners as a result of the lower rents achieved due to this burden.

Actually, that should read "all of which" and not "some of which", but there you go.

* Spotted by Thomas B Hall.